Life Cycle
Phase 1. seeds on forest floor
Phase 2. seedling establishment
Phase 3. growth and development
Phase 4. seed production and dispersal
Seeds on the Forest Floor
Germination:
- imbibition
- activation of metabolic processes
- embryo growth
Seed dormancy is defined as the failure of the seed to germinate under
favorable conditions. This trait is advantageous for seeds disseminated in
fall, as they do not germinate immediately and are held over until the spring.
The cold of winter could kill seedlings resulting from fall germination.
- Physiological Dormancy - internal, metabolic changes must occur
- Physical Dormancy - seed coat may be impermeable to water and gases or be
mechanically constraining
- Dormancy can be removed by stratification (cool moist treatment for
physiological dormancy) or scarification (reduce seed barrier by abrasion
for physical dormancy).
Vegetative Reproduction - root-collar - lignotuber - root sprouts -
fragmentation - stolons, runner, layering, tipping
Seedling Establishment
Pioneer species - bare areas, loblolly pine
Gap-phase species - under closed canopies and later released by opening of a
gap, yellow birch
Growth and Development
Primary growth - apical meristems at tips of roots and stems, growth in
length
- fixed growth: intermittent, resting bud, initiation in bud and elongation
separated in time
- free growth: continuous, initiation and elongation concurrent
Secondary growth - lateral meristems, cambium, growth in girth, growth
distribution and periodicity
Tree form
- excurrent, conical; conifers in northern and alpine
- decurrent, deliquescent; deciduous, broadleaf, southern pines
Seed Production and Dispersal
Flowering characteristics of common tree species
Age at which forest trees begin producing abundant seed crops
Typical developmental period required to produce mature seed in common tree
species
Phenology of seed production in common conifers
Periodicity in production of good seed crops in common tree species
Factors influencing periodicity in seed crops:
Site - earlier, heavier and
more consistent in southern part of range - dry south exposure, early seed
production
,- poor sites seem to stimulate early seed production
Climate - strong relationships are not clear
Insects - effects most noticeable in light seed years
Disease
Birds and mammals
Physiological
Seed Dispersal
wind, water, animals - see table of major dispersal types
most seeds are dispersed close to parent - 40 to 50 meters; advantage =
local adaptation, disadvantage = growing in competition with parent